Beetles of eastern North America

Preface 7  Acknowledgments 8  How to Use This Book 9  Classification 9  Key to Families 9  Family Diagnoses 9  Species Accounts 9  Species Identification 10   Introduction to Beetles 11  Beetle Anatomy 11  Behavior and Natural History 19  When and Where to Find Beetles 33  Observing and Photographing Beetles 36  Beetle Conservation and the Ethics of Collecting 38  Collecting and Preserving Beetles 39  Making a Beetle Collection 45  Keeping and Rearing Beetles in Captivity 49  Taking an Active Role in Beetle Research 52   Illustrated Key to the Common Beetle Families of Eastern North America 53  Beetles of Eastern North America 59  Reticulated beetles (Cupedidae) 60  Telephone-pole beetles (Micromalthidae) 61  Minute bog beetles (Sphaeriusidae) 62  Ground, tiger, and wrinkled bark beetles (Carabidae) 63  Whirligig beetles (Gyrinidae) 94  Crawling water beetles (Haliplidae) 96  Burrowing water beetles (Noteridae) 97  Predaceous diving beetles (Dytiscidae) 99  Water scavenger beetles (Hydrophilidae) 105  Clown beetles (Histeridae) 110  Minute moss beetles (Hydraenidae) 114  Featherwing beetles (Ptiliidae) 115  Primitive carrion beetles (Agyrtidae) 117  Round fungus beetles (Leiodidae) 118  Burying and carrion beetles (Silphidae) 120  Rove beetles (Staphylinidae) 124  Stag beetles (Lucanidae) 142  Bess beetles (Passalidae) 145  Enigmatic scarab beetles (Glaresidae) 146  Hide beetles (Trogidae) 147  Earth-boring scarab beetles (Geotrupidae) 149  Sand-loving scarab beetles (Ochodaeidae) 152  Scavenger and pill scarab beetles (Hybosoridae) 153  Bumble bee scarabs (Glaphyridae) 155  Scarab beetles (Scarabaeidae) 156  Plate-thigh beetles (Eucinetidae) 178  Minute beetles (Clambidae) 179  Marsh beetles (Scirtidae) 180  Cicada parasite beetles (Rhipiceridae) 183  Metallic wood-boring or jewel beetles (Buprestidae) 184  Pill or moss beetles (Byrrhidae) 195  Riffle beetles (Elmidae) 196  Long-toed water beetles (Dryopidae) 198  Travertine beetles (Lutrochidae) 200  Minute marsh-loving beetles (Limnichidae) 201  Variegated mud-loving beetles (Heteroceridae) 202  Water penny beetles (Psephenidae) 203  Ptilodactylid beetles (Ptilodactylidae) 204  Chelonariid beetles (Chelonariidae) 206  Callirhipid beetles (Callirhipidae) 207  Artematopodid beetles (Artematopodidae) 208  Rare click beetles (Cerophytidae) 209  False click beetles (Eucnemidae) 210  Throscid beetles (Throscidae) 211  Click beetles (Elateridae) 213  Net-winged beetles (Lycidae) 229  Glowworms (Phengodidae) 233  Fireflies, lightningbugs, and glowworms (Lampyridae) 234  False soldier and false firefly beetles (Omethidae) 237  Soldier beetles (Cantharidae) 238  Tooth-neck fungus beetles (Derodontidae) 243  Wounded-tree beetles (Nosodendridae) 244  Jacobsoniid beetles (Jacobsoniidae) 245  Skin beetles (Dermestidae) 246  Endecatomid beetles (Endecatomidae) 249  Bostrichid beetles (Bostrichidae) 250  Death-watch and spider beetles (Ptinidae) 252  Ship-timber beetles (Lymexylidae) 258  Bark-gnawing beetles and cadelles (Trogossitidae) 259  Checkered beetles (Cleridae) 263  Soft-winged flower beetles (Melyridae) 271  Fruitworm beetles (Byturidae) 274  Cryptic slime mold beetles (Sphindidae) 275  False skin beetles (Biphyllidae) 276  Pleasing fungus and lizard beetles (Erotylidae) 277  Root-eating beetles (Monotomidae) 281  Silken fungus beetles (Cryptophagidae) 283  Silvanid flat bark beetles (Silvanidae) 285  Flat bark beetles (Cucujidae) 288  Parasitic flat bark beetles (Passandridae) 289  Shining flower and shining mold beetles (Phalacridae) 290  Lined flat bark beetles (Laemophloeidae) 291  Short-winged flower beetles (Kateretidae) 293  Sap beetles (Nitidulidae) 295  Cybocephalid beetles (Cybocephalidae) 304  Palmetto beetles (Smicripidae) 305  Bothriderid beetles (Bothrideridae) 305  Minute bark beetles (Cerylonidae) 307  Handsome fungus beetles (Endomychidae) 308  Lady beetles (Coccinellidae) 311  Minute hooded and fungus beetles (Corylophidae) 320  Minute brown scavenger beetles (Latridiidae) 322  Hairy fungus beetles (Mycetophagidae) 323  Archeocrypticid beetles (Archeocrypticidae) 325  Minute tree-fungus beetles (Ciidae) 326  Polypore fungus beetles (Tetratomidae) 327  False darkling beetles (Melandryidae) 329  Tumbling flower beetles (Mordellidae) 333  Ripiphorid beetles (Ripiphoridae) 338  Zopherid beetles (Zopheridae) 340  Darkling beetles (Tenebrionidae) 344  Synchroa bark beetles (Synchroidae) 359  False longhorn beetles (Stenotrachelidae) 360  False blister beetles (Oedemeridae) 362  Blister beetles (Meloidae) 365  Palm and flower beetles (Mycteridae) 369  Conifer bark beetles (Boridae) 371  Dead log beetles (Pythidae) 372  Fire-colored beetles (Pyrochroidae) 373  Narrow-waisted bark beetles (Salpingidae) 376  Antlike flower beetles (Anthicidae) 377  Ischaliid beetles (Ischaliidae) 382  Antlike leaf beetles (Aderidae) 382  False flower beetles (Scraptiidae) 384  Disteniid longhorn beetles (Disteniidae) 387  Longhorn beetles (Cerambycidae) 388  Megalopodid leaf beetles (Megalopodidae) 428  Orsodacnid leaf beetles (Orsodacnidae) 429  Leaf and seed beetles (Chrysomelidae) 429  Pine flower snout beetles (Nemonychidae) 457  Fungus weevils (Anthribidae) 458  Cycad weevils (Belidae) 462  Leaf-rolling and thief weevils, and toothnose snout beetles (Attelabidae) 462  Straight-snouted and pear-shaped weevils (Brentidae) 466  Weevils, and snout, bark and ambrosia beetles (Curculionidae) 469   Appendix: Classification of the Beetles Covered in This Book 501  Glossary 523  Selected References and Resources 527  Photo and Illustration Credits 530  Index 537